Which Came First, the Fire-prone Habitat or the Fire-adapted Trait?

Just like soil and climate, fire has been shaping plant communities in fire-prone ecosystems around the world for millions of years. The proof is in the evolution of fire-adapted plant traits, a common theme for the following two research papers.

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Lamont, Byron B., and Tianhau He. 2016. Fire-proneness as a prerequisite for the evolution of fire-adapted traits. Trends in Plant Science 1493. 11pp. http://dx.doi.org/10.1016/j.tplants.2016.11.004